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THE TETRAD GRADUATE PROGRAM OFFERS DIVERSE TRAINING IN:

Biochemistry & Molecular Biology, Cell Biology,
Developmental Biology, & Genetics.

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About Us

The Tetrad graduate program offers diverse training and opportunities in four major research areas: biochemistry and molecular biology, cell biology, developmental biology, and genetics. The program is interdisciplinary, with an emphasis on collaborations among laboratories to solve outstanding problems in modern biology.

Program

The academic program for the first year includes three laboratory rotations as well as a core curriculum designed to provide students with a foundation for understanding and conducting research in experimental biology. All students take the four core courses during the first two quarters of the first year, and in the third quarter they choose a sequence of three intensive 2-week minicourses.
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Admissions Process

The Tetrad program is one of several UCSF graduate programs. You may apply to only one program – it is important to consider which program suits your interests and apply to only that program.

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Announcements

2017 NSF GRFP Fellowships awarded
Bushra Bibi
Antonio Cuevas-Navarro
Veronica Escalante
Irene Lui
Taylor Skokan
Chris Waters

2017 Harold M. Weintraub Graduate Student Award
to recognize outstanding achievement in Graduate Studies
awarded to Kelsie Eichel

Recent Publication

Larson AG, Elnatan D, Keenen MM, Trnka MJ, Johnston JB, Burlingame AL, Agard DA, Redding S, Narlikar GJ. Liquid droplet formation by HP1α suggests a role for phase separation in heterochromatin.. Nature. 2017 Jul 13;547(7662):236-240.