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Biochemistry & Molecular Biology, Cell Biology,
Developmental Biology, & Genetics.

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About Us

The Tetrad graduate program offers diverse training and opportunities in four major research areas: biochemistry and molecular biology, cell biology, developmental biology, and genetics. The program is interdisciplinary, with an emphasis on collaborations among laboratories to solve outstanding problems in modern biology.


The academic program for the first year includes three laboratory rotations as well as a core curriculum designed to provide students with a foundation for understanding and conducting research in experimental biology. All students take the four core courses during the first two quarters of the first year, and in the third quarter they choose a sequence of three intensive 2-week minicourses.
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Admissions Process

The Tetrad program is one of several UCSF graduate programs. You may apply to only one program – it is important to consider which program suits your interests and apply to only that program.

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2016 NSF GRFP Fellowships awarded

Kelsey Haas
Nairi Hartooni
Isabel Johnson

2016 Harold M. Weintraub Graduate Student Award
to recognize outstanding achievement in Graduate Studies
awarded to Trevor Sorrells

2016 Paul & Daisy Soros Fellowship Awarded to Nairi Hartooni

Recent Publication

Dumesic PA, Homer CM, Moresco JJ, Pack LR, Shanle EK, Coyle SM, Strahl BD, Fujimori DG, Yates JR, Madhani HD. Product binding enforces the genomic specificity of a yeast polycomb repressive complex. Cell. 2015 Jan 15;160(1-2):204-18. PMCID: PMC4303595.