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THE TETRAD GRADUATE PROGRAM OFFERS DIVERSE TRAINING IN:

Biochemistry & Molecular Biology, Cell Biology,Developmental Biology, & Genetics.

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About Us

The Tetrad graduate program offers diverse training and opportunities in four major research areas: biochemistry and molecular biology, cell biology, developmental biology, and genetics. The program is interdisciplinary, with an emphasis on collaborations among laboratories to solve outstanding problems in modern biology.

Program

The academic program for the first year includes three laboratory rotations as well as a core curriculum designed to provide students with a foundation for understanding and conducting research in experimental biology. All students take the four core courses during the first two quarters of the first year, and in the third quarter they choose a sequence of three intensive 2-week minicourses.
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Admissions Process

The Tetrad program is one of several UCSF graduate programs. You may apply to only one program – it is important to consider which program suits your interests and apply to only that program.

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Announcements

2020 NSF GRFP Fellowship awarded
 to:

Wei Gordon

Seesha Takagishi

Connie Yan

2020 NASEM Ford Foundation Fellowship awarded to:

Evelyn Hernandez

Eric Simental

Congratulations to all of our awardees!

Recent Publication

Mendoza SD, Nieweglowska ES, Govindarajan S, Leon LM, Berry JD, Tiwari A, Chaikeeratisak V, Pogliano J, Agard DA, Bondy-Denomy J. A bacteriophage nucleus-like compartment shields DNA from CRISPR nucleases. Nature. 2020 Jan;577(7789):244-248. doi: 10.1038/s41586-019-1786-y. Epub 2019 Dec 9.