Tetrad Graduate Program

World-Renowned Collaborative Spirit

UCSF graduate programs were among the first in the country to break down traditional barriers between departments and disciplines, and interdisciplinary thinking permeates the program curriculum and faculty.

Highly Innovative and Constantly Evolving Curriculum

Interactive, discussion-based classes complement hands-on research projects.

Curiosity-Driven Research

Faculty and students in the Tetrad Graduate Program are driven by curiosity, leading to fundamental discoveries in all areas of basic biology and biomedical sciences.

Recent Awards

2026 NSF GRFP Fellowship awarded  to:

Gabriela Canales

Maia Kirkegaard

Eileen Tzng

Congratulations to the Tetrad student awardees!

Recent Publications

Howard MK, Hoppe N, Huang XP, Mitrovic D, Billesbølle, CB, Macdonald CB, Mehrotra E, Rockefeller Grimes P, Trinidad, DD, Delemotte L, English JG, Coyote-Maestas W, & Manglik A
Molecular basis of proton sensing by G protein-coupled receptors.
Cell vol. 188,3 (2025): 671-687.e20. doi:10.1016/j.cell.2024.11.036.
Estevam, GO, Linossi EM, Macdonald CB, Espinoza CA, Michaud JM, Coyote-Maestas W, Collisson EA, Jura N, & Fraser JS
Conserved regulatory motifs in the juxtamembrane domain and kinase N-lobe revealed through deep mutational scanning of the MET receptor tyrosine kinase domain.
eLife vol. 12 RP91619. 13 Sep. 2024, doi:10.7554/eLife.91619.