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THE TETRAD GRADUATE PROGRAM OFFERS DIVERSE TRAINING IN:

Biochemistry & Molecular Biology, Cell Biology,
Developmental Biology, & Genetics.

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About Us

The Tetrad graduate program offers diverse training and opportunities in four major research areas: biochemistry and molecular biology, cell biology, developmental biology, and genetics. The program is interdisciplinary, with an emphasis on collaborations among laboratories to solve outstanding problems in modern biology.

Program

The academic program for the first year includes three laboratory rotations as well as a core curriculum designed to provide students with a foundation for understanding and conducting research in experimental biology. All students take the four core courses during the first two quarters of the first year, and in the third quarter they choose a sequence of three intensive 2-week minicourses.
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Admissions Process

The Tetrad program is one of several UCSF graduate programs. You may apply to only one program – it is important to consider which program suits your interests and apply to only that program.

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Announcements

2019 NSF GRFP Fellowship awarded
 to

Amanda Chung

Congratulations, Amanda!

Recent Publication

Dalal CK, Zuleta IA, Lohse MB, Zordan RE, El-Samad H, Johnson AD. A population shift between two heritable cell types of the pathogen Candida albicans is based both on switching and selective proliferation. Proc Natl Acad Sci U S A. 2019 Dec 10. pii: 201908986. doi: 10.1073/pnas.1908986116. [Epub ahead of print].